Beloved Galápagos tortoise Gramma, a longtime favorite at the San Diego Zoo, has died at an estimated age of 141.

Gramma, one of the San Diego Zoo’s most beloved residents and among the oldest Galápagos tortoises in human care, has died at an estimated age of 141. Her extraordinary lifespan stretched across generations, world events, and countless visitors who came to admire her gentle presence. Born in the late 1800s, Gramma lived through decades of conservation work and helped educate millions of guests about the species’ history and fragility. Her passing marks the end of an era for the zoo and for admirers around the world.








